My observation was regarding the resultant force exerted by the explosion. The compression ratio will remain the same if the volume at the cillynder cavity is unchanged.Again, I don't disagree that a change in compression would cause artifacts in addition to the spark placement, but how are you inferring that the compression is lower with the acrylic head?
But think also about this: air behaves pretty much like a spring when it's compressed. And an assimetric compression, such as the one present in the cillynder shown in the video, will leave zones within its volume with significant pressure differences. This will affect the uniformity at which combustion takes place.