No, the 3V are likely just at the edge of lighting the LEDs at all. As the cells deplete, the voltage will drop quickly to a point where they are too dim, long before they are completely depleted. For example, the 3V battery will be about 2.3V when half depleted, a that probably will not light the lights at all.I assume two the the 2 C cell packs in parallel for 3v would be the best way to go for longevity? (in the photo they are in series for 6v)
3 cells at 4.5V (with different resistors) would likely give you better runtime than 4 cells at 3V. The 4.5V battery will still be around 3V when fully depleted and the lights will still light, though dimmer because of the higher resistors.



